Systems and methods are provided for advancing a fuselage section of an aircraft. One embodiment is a system that includes a series of plates configured to be sequentially affixed along a length of the fuselage section, and a track configured to form a frictional fit with the plates. The track includes drive units configured to form nips retaining the series of plates, and that are configured to advance the fuselage section along the track in a process direction, and indexing elements that are configured to engage with indexing elements at the plates during pauses between operation of the drive units. The system also includes tools configured to perform work on the fuselage section while the indexing elements of the track are engaged.